Background Info

For more information about the Clover Project and how you can use it in your classroom, Click <Here> and Dr. Kenneth Olsen will explain some of the science behind his research with white clover at Washington University in St. Louis, Missouri, and how it can benefit you and your students.

Some useful background information students may need in order to complete the cyanogenesis assay are found below:

What students need to know about white clover before getting started: Background information for teachers

Beneficial vocabulary

Facts about evolution, natural selection, and adaptive polymorphism in white clover.


The creators of the modified clover cyanogenesis lab protocol are high school biology teachers Joy Auclair and Anne Puzzo of Granby High School in Conneticut.

The lab protocol was piloted by high school biology teachers Chuck McWilliams (Maplewood Richmond Heights School District, St. Louis, MO) and David Ganey (Jefferson City High School, Jefferson City, MO).

During the summers of 2010 and 2011, this website and accompanying materials were created by Sheryl Silverberg, a high school biology teacher for the Lindbergh School District in St. Louis, Missouri, and an intern working for Dr. Olsen.

In the fall of 2011, Chuck McWilliams revised this website.  Linda Small made additional revisions in the fall of 2018.

Linda Small currently manages the project.

Note: All of the teachers listed above have made valuable contributions to this program and are graduates of the "Life Science for a Global Community" graduate program for teachers in the College of Arts and Sciences at Washington University in St. Louis, MO.