Entrainment is the process by which an oscillator synchronizes to a periodic input, like how we sync to the daily light-dark cycle. This tutorial, aimed at students of biological timing who wish to explore the various models of how circadian pacemakers entrain to daily cues, starts with classic readings by Colin Pittendrigh. We will meet in the same place and time as Clocksclub.


  1. Phase repsonse curves: pedictive value
  2. Dawn to dusk and jet lag: Evidence for parametric vs non-parametric entrainment
  3. A clock for all seasons: Coiupled (E&M) oscillators/files/pages/imce/photo.synth.bio/r03.pdf
  4. Limit cylcel oscillators